LINNEAN SOCIETY 的动物学杂志发表了关于系统和进化动物学以及与这些领域相关的比较、功能和其他研究的论文。包括对已灭绝和现存动物的研究。评论也发表；这些可能会受到编辑委员会的邀请，但也可能会考虑不请自来的评论。动物学杂志在动物学家中也有广泛的流通，虽然不排除狭隘的专业论文，但潜在的作者应该牢记这一读者群。

The Zoological Journal of the Linnean Society is a zoological journal published by the LINNEAN SOCIETY, primarily focusing on research papers related to systematic and evolutionary zoology, as well as comparative, functional, and other studies associated with these fields. The journal includes studies on both extinct and extant animals. Additionally, it publishes review articles, which may be invited by the editorial board or considered from unsolicited submissions. The journal enjoys a broad circulation among zoologists, and while it does not exclude niche professional papers, potential authors should keep in mind this diverse readership.

《Zoological Journal of the Linnean Society》是LINNEAN SOCIETY出版的动物学期刊，主要发表关于系统与进化动物学的研究论文，以及与这些领域相关的比较、功能和其他研究。期刊内容涵盖对已灭绝和现存动物的研究。此外，期刊也会发表评论文章，这些评论可能是编辑委员会邀请的，也可能是自荐的。该期刊在动物学界有广泛的影响力，尽管不排除狭隘的专业论文，但潜在的作者应考虑到这一读者群的广泛性。

Zoological Journal of the Linnean Society

New data are presented on the neurocranial complex (endocranium plus intimately associated bones of the palate) of the Late Devonian (Famennian) ‘rhynchodipterid’ lungfish Soederberghia groenlandica from the Aina Dal Formation (Celsius Bjerg Group) of East Greenland. Only the otic and occipital regions of the braincase are ossified in Soederberghia. The neurocranium of this genus shares a series of derived features with ‘Griphognathus’ whitei, including a cranial centrum, fenestrate lateral cristae, and gutters on the ventral surface of the parasphenoid interpreted as accommodating the lateral dorsal aortae. The interrelationships of early lungfishes have been the subject of considerable disagreement. New data from Soederberghia are coupled with a data set focused on the character-rich neurocranial complex in order to examine the systematic utility of this underexploited morphological system. Different methods of phylogenetic inference (maximum parsimony, Bayesian) return broadly consistent results. The Early to Middle Devonian forms Dipnorhynchus, Stomiahykus and Uranolophus are placed among the most basal of lungfishes. ‘Holodontids’ plus ‘rhynchodipterids’ (comprising Griphognathus and Soederberghia) occupy an apical position, and are separated from the earliest lungfishes by a paraphyletic assemblage of taxa generally identified as ‘chirodipterids’ and ‘dipterids.’ This finds broad agreement with the results of previous cladistic studies focused on non-neurocranial data sets, but diverges from functional-adaptive scenarios that posit three lineages of early lungfishes based on aspects of the dentition. As currently defined, both ‘chirodipterids’ and the genus Chirodipterus are heterogeneous assemblages; this analysis fails to find support for the monophyly of either. While this study indicates that Griphognathus is probably paraphyletic, it nevertheless supports a close relationship between the nominal species of this genus and Soederberghia.

MATT FRIEDMAN

The interrelationships of Devonian lungfishes (Sarcopterygii: Dipnoi) as inferred from neurocranial evidence and new data from the genus Soederberghia Lehman, 1959

Sauropoda is among the most diverse and widespread dinosaurlineages, having attained a near-global distribution by the Middle Jurassic that was built on throughout the Cretaceous. These giganticherbivores are characterized by numerous skeletal specializationsthat accrued over a 140 million-year history. This fascinating evolutionaryhistory has fuelled interest for more than a century, yet aspects of sauropod interrelationships remain unresolved. This paper presentsa lower-level phylogenetic analysis of Sauropoda in two parts. First, the two most comprehensive analyses of Sauropoda are critiqued toidentify points of agreement and difference and to create a core of character data for subsequent analyses. Second, a generic-levelphylogenetic analysis of 234 characters in 27 sauropod taxa is presentedthat identifies well supported nodes as well as areas of poorerresolution. The analysis resolves six sauropod outgroups to Neosauropoda, which comprises the large-nostrilled clade Macronaria and the peg-toothedclade Diplodocoidea. Diplodocoidea includes Rebbachisauridae, Dicraeosauridae, and Diplodocidae, whose monophyly and interrelationships are supportedlargely by cranial and vertebral synapomorphies. In contrast, thearrangement of macronarians, particularly those of titanosaurs, are based on a preponderance of appendicular synapomorphies. The purported Chinese clade 'Euhelopodidae' is shown to comprisea polyphyletic array of basal sauropods and neosauropods. The synapomorphiessupporting this topology allow more specific determination for themore than 50 fragmentary sauropod taxa not included in this analysis. Their distribution and phylogenetic affinities underscore the diversity of Titanosauria and the paucity of Late Triassic and Early Jurassicgenera. The diversification of Titanosauria during the Cretaceousand origin of the sauropod body plan duringthe Late Triassic remain frontiers for future studies.

Jeffrey A. Wilson

Sauropod dinosaur phylogeny: critique and cladistic analysis

Abstract Dryinus is a cosmopolitan genus and the most diverse group within Dryininae. Herein, we present the first phylogenetic analyses for species groups of Dryinus from the Neotropical region, including three extant genera of Dryininae, one fossil, and a rich taxon sampling of extant species. A total of 116 characters were coded to 42 terminal taxa, including 28 species representing Dryinus of the Neotropics. Our analyses showed that Dryinus is paraphyletic, with the two species D. caraibicus (Olmi) and D. lamellatus (Olmi) coming out separately from the remaining species. A new genus, Dirrhinodryinus, is here described to accommodate them. Based on the phylogeny, Dryinus s.s. is divided into six species groups: autumnalis, catarinae, constans, fiorii, ruficeps, and magnificus. Of the four previously recognized species groups, only two were recovered here: autumnalis and constans. Additionally, we corroborated the synonym of Megadryinus under Dryinus. Considering the evidence that the Neotropical Dryinus form an independent clade, our expectation is that the new classification proposed here will not be applicable to the fauna of other biogeographic regions.

André L Martins

Gabriel A R Melo

Phylogenetic analysis of Neotropical <i>Dryinus</i> using morphological characters: new proposed classification for species groups and description of a new genus (Hymenoptera: Dryinidae)

利用形态特征对新热带区<i>Dryinus</i>进行系统发育分析：物种群的新分类建议及新属描述（膜翅目：蚁蜂科）

Abstract Species in the beetle family Histeridae exhibit numerous defensive adaptations, from a generally compact and spheroid body shape to retractable tarsi, legs, heads, and antennae. Modes of antennal concealment are particularly variable, underscoring the importance of protecting these essential sensory structures. Understanding the evolution of these morphological features has been hindered by a weak understanding of the family’s basal phylogeny, and corresponding patterns of niche-use and morphological evolution. Cretaceous amber fossils have been proliferating, filling in important morphological gaps right around the time of the family’s explosive radiation. Here we describe 10 new species from Cretaceous Burmese ambers, six of them representing new genera: Palaeabraeus glabrus gen. nov. et sp. nov. (Abraeinae); Pseudacritus extinctus gen. nov. et sp. nov. (Abraeinae); Miculissima excavata gen. nov. et sp. nov. (unplaced to subfamily); Placatister cascus gen. nov. et sp. nov. (unplaced to subfamily); Cretomalus tibiodentatus gen. nov. et sp. nov. (Dendrophilinae: Paromalini); Prodigister tricostatus gen. nov. et sp. nov. (unplaced to subfamily); Anapleus punctulatus sp. nov. (Dendrophilinae: Anapleini); Pantostictus hirsutus sp. nov. (Abraeinae: Pantostictini); Phasmister kraliceki sp. nov. (Onthophilinae); and Phasmister planatus sp. nov. (Onthophilinae). A revised phylogenetic analysis that includes these taxa supports previous hints that inquilinous forms were prevalent and diverse during the Cretaceous. These results also support the contention that pressures in these environments to protect the critically important sensory apparatus, the antenna, were strong, driving multiple origins of two of the more protective configurations among extant taxa. Some of the new genera remain unplaced to subfamily, underscoring the gaps that still remain in the early fossil record for the family, and the continued need for more comprehensive analyses of combined data from extant taxa, along with additional extinct forms that we may assume will continue to be discovered.

Jan Simon-Pražák

Shûhei Yamamoto

Tomáš Lackner

Martin Fikáček

Jakub Prokop

Michael S Caterino

A bonanza of Cretaceous fossils provides insights into the evolution of antennal protection in clown beetles (Coleoptera: Histeridae)

白垩纪化石的丰富发现揭示了小丑甲虫（鞘翅目：隐翅虫科）触角保护的演化

Abstract We here describe Terastiodontosaurus marcelosanchezi, a new amphisbaenian genus and species from the Eocene of Chambi, Tunisia. Using micro-computed tomography (μCT), we document the peculiar anatomy of the new taxon, which is characterized by extreme dental morphology, including one massive tooth on the maxilla and dentary, flat cheek teeth, and an array of other diagnostic features that readily differentiate it from all other amphisbaenians. We also redescribe the oldest named African amphisbaenian, Todrasaurus gheerbranti, from the late Palaeocene of Morocco, using μCT. Phylogenetic analysis recovers Terastiodontosaurus and Todrasaurus as sister taxa and provides strong support for a sister-group relationship of those two large-toothed amphisbaenians with extant Trogonophis. Accordingly, Todrasaurus shows that the divergence of crown Trogonophidae occurred much earlier than currently thought. Our survey of μCT scans reveals that Terastiodontosaurus, Todrasaurus, and Trogonophis are characterized by a great enamel thickness on their teeth, a feature that is absent in other examined amphisbaenians. Size estimates show that Terastiodontosaurus was the largest known amphisbaenian ever to have lived, with an estimated skull length of &amp;gt;5 cm. Based on new muscle data of Trogonophis, we estimate very high bite forces for Terastiodontosaurus, which would allow it to crush a wide variety of snails.

Georgios L Georgalis

Krister T Smith

Laurent Marivaux

Anthony Herrel

El Mabrouk Essid

Hayet Khayati Ammar

Wissem Marzougui

Rim Temani

Rodolphe Tabuce

The world’s largest worm lizard: a new giant trogonophid (Squamata: Amphisbaenia) with extreme dental adaptations from the Eocene of Chambi, Tunisia

世界最大的蠕蜥：来自突尼斯沙姆比始新世的巨型托戈诺菲德（有鳞目：双足蜥科）及其极端牙齿适应性

Abstract Fish skulls are often highly kinetic, with multiple linkage and lever systems powered by a diverse suite of muscles. Comparative analysis of the evolution of soft-tissue structures in the fish skull is often limited under traditional approaches, while new imaging techniques like diceCT (diffusible iodine-based contrast-enhanced computed tomography) allow for high-resolution imaging of muscles in situ. Darters (Percidae: Etheostomatinae) are a diminutive and species-rich clade of lotic freshwater fishes, which show diverse head shapes believed to be associated with different foraging strategies. We used diceCT to sample all major cranial adductors and abductors responsible for movement of the jaw, hyoid, operculum, and suspensorium from 29 species. We applied comparative phylogenetic approaches to analyse the evolutionary trends in muscle size across the clade. We found two major patterns: (i) darter cranial muscles show fundamental trade-offs relating to investment in musculature, as well as buccal expansion vs. biting attributes; early divergence in muscle size appears to be associated with shifts in habitat use and foraging; (ii) darter adductor mandibulae show high variation in architecture (fibre orientation, divisions). This study highlights how new imaging techniques can provide novel insights into the anatomy of even well-sampled/represented clades.

J H Arbour

S Ramazan

S Clark

Adaptation and innovation in darter fish cranial musculature (Etheostomatinae: Percidae): insights from diceCT

达特鱼颅骨肌肉的适应与创新（Etheostomatinae: Percidae）：基于diceCT的见解

Abstract Research interest in the diversity and evolutionary history of herring-like fossils (subcohort Clupei) has increased in recent decades. However, little is known about the relationships between fossils assigned to Clupei, particularly those that are demonstrably related to extant herring-like members of the order Clupeiformes. To help bridge this gap, we present a new morphological phylogeny that includes representatives of all major clupeiform lineages. The data matrix consists of 192 characters, drawn from 79 extant and 37 fossil taxa and selected to be readily identifiable in fossils. Most clupeiform families are recovered as monophyletic, and their interrelationships are generally compatible with previous morphological hypotheses. The phylogenetic positions of six fossil clupeiform taxa are resolved, as the results of all analyses (unconstrained/constrained parsimony and Bayesian inference) are consistent. Twenty-one fossil taxa are incertae sedis, including two species of Ellimmichtyiformes, which are recovered as Clupeoidei in the parsimony analyses. The relationships of the rest of the fossils are interpreted with varying degrees of certainty. The evolution of key morphological characters is discussed in light of the new results, and four fossil taxa are proposed as calibration points for future dating studies.

Charalampos Kevrekidis

Timo Moritz

Alexander F Cerwenka

Elena Bauer

Bettina Reichenbacher

Uncovering the relationships among herring-like fossils (Clupei: Teleostei): a phylogenetic analysis

揭示鲱鱼类化石（鲱形目：真骨鱼类）之间的亲缘关系：系统发育分析

Abstract About 300 valid species are classified within the order Rajiformes, the largest group of chondrichthyans. The monophyly of this order is highly supported by morphological and molecular inferences, but little is known about the mandibular, hyoid, and hypobranchial muscles of its members. This study aims to describe and illustrate the morphological variation in mandibular, hyoid, and hypobranchial muscles among rajiforms, review their terminology and discuss the phylogenetic and functional implications of conditions observed. Twenty-two ethanol-preserved specimens belonging to 19 of the 36 skate genera currently considered valid were manually dissected. Nine new characters are proposed and tested cladistically. The main differences observed are the relative width of muscle units of the levator palatoquadrati, insertion point and extension of the intermanibularis posterior, relative position of the facialis nerve and the levator hyomandibularis muscle, origin and insertion of the coracomandibularis, extension of the coracohyomandibularis, and occurrence of the preorbitalis medialis and intermandibularis anterior. When analysed together with other morphological features, muscle characters have shown to play a key role in the understanding of interrelationships among skates. Nevertheless, additional studies are needed to improve our knowledge of rajiform systematics and the functionality of mandibular, hyoid, and hypobranchial muscles.

Abbreviation	Zool. J. Linn. Soc.
Journal Impact	2.92
Quartiles(Global)	ZOOLOGY(Q1)

ISSN	0024-4082, 1096-3642
h-index	95

Publication Information	Publisher: Oxford University Press，Publishing cycle: Monthly，Journal Type: journal，Open Access Journals: No
Basic data	Year of publication: 1949，Proportion of original research papers: 95.68%，Self Citation Rate:6.70%， Gold OA Rate: 16.18%
Average review cycle	网友分享经验：较慢,6-12周
Average recruitment ratio	网友分享经验：较易

Zoological Journal of the Linnean Society

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The interrelationships of Devonian lungfishes (Sarcopterygii: Dipnoi) as inferred from neurocranial evidence and new data from the genus Soederberghia Lehman, 1959

Sauropod dinosaur phylogeny: critique and cladistic analysis

Phylogenetic analysis of Neotropical Dryinus using morphological characters: new proposed classification for species groups and description of a new genus (Hymenoptera: Dryinidae)

A bonanza of Cretaceous fossils provides insights into the evolution of antennal protection in clown beetles (Coleoptera: Histeridae)

The world’s largest worm lizard: a new giant trogonophid (Squamata: Amphisbaenia) with extreme dental adaptations from the Eocene of Chambi, Tunisia

A bonanza of Cretaceous fossils provides insights into the evolution of antennal protection in clown beetles (Coleoptera: Histeridae)

The world’s largest worm lizard: a new giant trogonophid (Squamata: Amphisbaenia) with extreme dental adaptations from the Eocene of Chambi, Tunisia

Adaptation and innovation in darter fish cranial musculature (Etheostomatinae: Percidae): insights from diceCT

Uncovering the relationships among herring-like fossils (Clupei: Teleostei): a phylogenetic analysis

Mandibular, hyoid, and hypobranchial musculature in skates (Chondrichthyes: Batoidea: Rajiformes): revised terminology and phylogenetic implications